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NIH Training Grant Program

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Cell and Molecular Biology Training Program

We seek to train the next generation of researchers in cell and molecular biology with a deep understanding of core disciplines and the ability to plan important, incisive research and execute their plans with appropriate technology and methodology, from the classic to the cutting edge.

To join, apply via one of the following Graduate Options: Biology, BMB, Chemistry, Bioengineering, Geobiology, Computer Science, Neurobiology, CNS.


Our training activities comprise eight main components described in detail below this overview:

  1. each student's individual research program, guided by one or more faculty members and carried out within a group of other students and postdoctoral fellows having related interests;
  2. a set of courses comprising of Topics in molecular and cell biology (Bi250a), a  class in reading, writing, reviewing, expiremental design and reproducibility (Bi253), three courses from a menu of relevant courses;
  3. option-specific preparation for qualifying or candidacy examinations;
  4. an extensive program of formal and informal seminars and research meetings;
  5. a course and discussion in responsible conduct of research (Bi252);
  6. a spring term research seminar during which students present their own work and discuss that of other students (Bi/BMB 251);
  7. yearly meetings with a thesis committee to provide an update on progress towards the Ph.D. degree; and
  8. experience in teaching through service as a teaching assistant.

Training Faculty by Division

David Anderson, BBE
Alexei Aravin, BBE
Frances Arnold, CCE
David Baltimore, BBE
Jacqueline Barton, CCE
Pamela Bjorkman, BBE
Marianne Bronner, BBE
Long Cai, BBE
Judith Campbell, BBE/CCE
David Chan, BBE
William Clemons, CCE
Peter Dervan, CCE
Michael Dickinson, BBE
Dennis Dougherty, CCE
Michael Elowitz, BBE/EAS

Lea Goentoro, BBE
Viviana Gradinaru, BBE
Mitchell Guttman, BBE


Bruce Hay, BBE
Andre Hoelz, CCE
Linda Hsieh-Wilson, CCE
Rustem Ismagilov, CCE
Grant Jensen, BBE
Jared Leadbetter, EAS
Henry Lester, BBE

Carlos Lois, BBE

Stephen Mayo, BE/CCE

Sarkis Mazmanian, BBE

Elliot Meyerowitz, BBE

Richard Murray, BBE/EAS

Dianne Newman, BBE

Alison Ondrus,CCE

Victoria Orphan, GPS

Lior Pachter, BBE

Joseph Parker, BBE

Robert Phillips, EAS
Niles Pierce, EAS

David Prober, BBE

LuLu Qian, BBE
Douglas Rees, CCE
Sarah Reisman, CCE
Ellen Rothenberg, BBE
Shu-Ou Shan, CCE
Mikhail Shapiro, CCE
Angelike Stathopoulos, BBE
Paul W. Sternberg*, BBE

Matthew Thomson, BBE

David Van Valen, BBE
Alex Varshavsky, BBE

Rebecca Voorhees, BBE
Zhen-Gang Wang, CCE
Barbara Wold, BBE
Kai Zinn, BBE


*Funded in part by National Institutes of Health, National Institute of General Medical Sciences grant 2T32GM007616 (P. W. Sternberg, PI).